(was emb*-8502; emb*-8503) embryo development blocked at late transition stage. Scutellum elongate, shoot apex poorly developed, embryo necrotic. Normal segregation ratio. (J. Clark). Allele emb*-8503 more variable in expression than emb*-8502. Genetic 2005 1 coordinates determined from these recombination and map data: emb4 TB-1Sb; see MaizeGDB ID:139767. This gene was originally curated in MaizeGDB Database and the map position was displayed on the maize bins map if feasible.