Awned indica cultivar Roti was crossed with awnless 'Club mutant' and awnless 'Dwarf mutant'. In Roti X Dwarf mutant, awned F1 and a 9-awned: 7 awnless segregating ratio in F2 indicate the interaction of two pairs of complementary genes, An6 (Ana) and An7 (Anb), which are carried by Roti and necessary for awn development. In the cross of Roti X Club mutant, Awnless F1 and a 9 awned: 247 awnless segregating ratio in F2 show that Club mutant carries two dominant inhibitors, An8 (IAna) and An9 (IAnb), which suppress the expression of An6 (Ana) and An7 (Anb).