Both NADH-and NADPH-dependent activities of nitrate reductase (NR) were considerably reduced, compared to the wild type. Xanthine dehydrogenase (XDH) activities and molybdenum cofactor biosynthesis abilities (Moco activities) were deficient. Nitrate reductase activities increased compared to the wild type. Similar to cnx1. Compared with cnx1, NADH-NR activity was not recovered while adding 0.5 mM molybdate to the growth medium.